About EFAs
EFAs and Disease
EFAs and Beauty
EFAs and Pets
EFA Glossary
FAQs
Supplement Sources
Selected References

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The information on essential fatty acids (EFAs) presented
in this website is based on documented scientific research.
This list summarizes a selection of references used
to compile the information on this site.
General
- al-Shabanah, OL. Effect of Evening Primrose Oil
on Gastric Ulceration and Secretion Induced by Various
Ulcerogenic and Necrotizing Agents in Rats, Food
Chemistry Toxicology, Vol. 35, p. 769, 1997.
- Bierenbaum ML, et al. Reducing atherogenic risk
in hyperlipemic humans with flax seed supplementation:
a preliminary report. Journal of the American College
of Nutrition, Vol. 12(5), pp. 501-4, 1993.
- Cunnane SC, et al. Nutritional attributes of traditional
flaxseed in healthy young adults. American Journal
of Clinical Nutrition. Vol. 61(1):62-68, 1995.
- Hamadeh MJ, et al. Nutritional aspects of flaxseed
in the human diet. Proceedings of the Flax Institute.
Vol. 4, pp 48-53, 1992.
- Harris W.S. n-3 Fatty acids and serum lipoproteins:
human studies. American Journal of Clinical Nutrition.
Vol. 65 (Supplement), pp 1645-54, 1997.
- Horrobin, D. Loss of delta-6-desaturase activity
as a key factor in aging. Medical Hypothesis,
Vol. 9, pp 1211-1220, 1981.
- Parbtani A and Clark WF. In: Flaxseed in Human Nutrition.
Cunnane SC and Thompson LU, eds. Champaign, IL: AOCS
Press, pp. 244-260, 1995.
- Zurier et al. Gamma-linolenic acid: Metabolism and
its roles in Nutrition and Medicine. pp. 129-136,
1996.
- Zurier RB, et al. Modification of human lymphocyte
and monocyte function by gamma-linolenic acid (GLA),
an unsaturated fatty acid. Studies in vitro and in
vivo. Proc Annual Meeting of the American Oil Chemists'
Society. p. 24, 1998
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Cardiovascular
Health
- Appel L.J., et al. Does Supplementation of Diet
with 'Fish Oil' Reduce Blood Pressure'? A Meta-analysis
of Controlled Clinical Trials. Archives of Internal
Medicine. Vol. 153 (12), pp 1429-38, 1993.
- Bang H.O., Dyerberg, J., Hjoorne, N. The composition
of food consumed by Greenland Eskimos. Acta Medical
Scandinavia. Vol. 200, pp 69-73, 1976.
- Bierenbaum ML, et al. Reducing atherogenic risk
in hyperlipemic humans with flax seedsupplementation:
a preliminary report. Journal of the American College
of Nutrition. Vol. 12(5), pp. 501-4, 1993.
- De Lorgeril et al. Mediterranean alpha-linolenic
acid-rich diet in secondary prevention of coronary
heart diseases. Lancet. Vol. 343, pp. 1454-1459, 1994.
- Hu FB, et al. Dietary intake of alpha-linolenic
acid and risk of fatal ischemic heart disease among
women American Journal of Clinical Nutrition,
Vol. 69, pp. 890-7, 1999.
- Jacob R. et al. Prolongation of life span in hypertensive
rats by dietary interventions. Effects of garlic and
linseed oil. Basic Research in Cardiology,
Vol. 92(4), pp 223-232, 1997.
- McLennan P., et al. The cardiovascular protective
role of docosahexaenoic acid. European Journal
of Pharmacology. Vol.300, pp 83-89, 1995.
- Renaud S, et al. Cretan Mediterranean diet for prevention
of coronary heart disease. American Society for
Clinical Nutrition. Vol. 61(Supplement), pp 1360-67.
1995.
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Joint
Health
- Ariza-Ariza, R. Omega-3 fatty acids in rheumatoid
arthritis: an overview. Seminars in Arthritis and
Rheumatism, Vol. 27, pp. 366-370, 1998.
- Belch JJ, et al. Effects of altering dietary essential
fatty acids on requirements for non-steroidal anti-inflammatory
drugs in patients with rheumatoid arthritis: a double-blind
placebo controlled study, Annals of Rheumatoid
Disease, Vol. 47, pp 96-104 1988.
- Blackburn GL. Nutrition and inflammatory events:
highly unsaturated fatty acids (omega-3 vs omega-6)
in surgical injury. Proceedings of the Society
for Experimental Biology and Medicine. Vol. 200,
pp 183-188, 1992.
- Calder P. Immunoregulatory and anti-inflammatory
effects of n-3 polyunsaturated fatty acids. Brazillian
Journal of Medical and Biological Research, Vol
31(4), pp. 467-90, 1998.
- Guillermo, Tate, et al. Suppression of Acute and
Chronic Inflammation by Dietary Gamma Linolenic Acid,
Journal of Rheumatology, Vol. 16, pp 729-33,
1989.
- Kremer, J.M. n-3 fatty acid supplements in rheumatoid
arthritis. American Journal of Clinical Nutrition,
Vol. 71(1), pp 349S-3451S, Jan. 2000.
- Leventhal, LJ, et al. Treatment of Rheumatoid Arthritis
with Gamma Linolenic Acid, Annals of Internal Medicine,
Vol. 119, pp 867-73, 1993.
- Pharmaceutical Information Associates, Nonprescription
NSAIDs: Efficacy and Safety, Medical Sciences Bulletin,
June 1994.
- Pullman-Mooar S et al. Alteration of the cellular
fatty acid profile and the production of eicosanoids
in human monocytes by gamma-linolenic acid. Arthritis
and Rheumatism, Vol. 33(10), pp 1526-33, Oct.
1990.
- Rothman et al. Effects of unsaturated fatty acids
on interleukin-1beta production by human monocytes.
Cytokine, Vol. 9(12), pp 1008, 1997.
- Zurier RB, et al. Gamma-Linolenic acid treatment
of rheumatoid arthritis. A randomized, placebo-controlled
trial. Arthritis Rheumatology, Vol. 39, No
11, pp 1808-17, 1996.
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Diabetes
- Cotter, C., et al. Metabolic and Vascular Factors
in the Pathogenesis of Diabetic Neuropathy, Diabetes,
Vol. 46, Suppl. 2, pp S31-S37, 1997.
- Horrobin DF. The effects of Gamma Linolenic Acid
on breast pain and diabetic neuropathy: possible non-eicosanoid
mechanisms.
Prostaglandins Leukotr Essential Fatty Acids,
Vol. 48, pp 101-104, 1993.
- Jamal GA, et al. Gamma Linolenic acid in diabetic
neuropathy. Lancet, Vol. 10. p. 1098, 1986.
- Keen, H, et al. Treatment of diabetic neuropathy
with gamma-linolenic acid. The gamma-Linolenic Acid
Multicenter Trial Group. Diabetes Care. Vol.
16, No. 1, pp 8-15, 1993.
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Skin
Health
- Andreassi, M., et al. Efficacy of Gamma Linolenic
Acid in the Treatment of Patients with Atopic Dermatitis,
The Journal of International Medical Research,
Vol. 25, pp. 266-74, 1997.
- Henz B.M., Double-blind, multicentre analysis of
the efficacy of borage oil in patients with atopic
eczema, British Journal of Dermatology, Vol.
140, pp. 685-88, 1999.
- Morse P.F., et al. Meta-analysis of placebo-controlled
studies of the efficacy of Epogam in the treatment
of atopic eczema. Relationship between plasma essential
fatty acid changes and clinical response, British
Journal of Dermatology, Vol. 121, pp. 75-90, 1989.
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Women's
Health
- Brush MG, Watson SJ, Horrobin DF, Manku MS. Abnormal
essential fatty acid levels in plasma of women with
premenstrual syndrome. American Journal of Obstetrics
and Gynecology. Vol. 150, pp 363-366, 1984.
- Horrobin DF. The effects of Gamma Linolenic Acid
on breast pain and diabetic neuropathy: possible non-eicosanoid
mechanisms. Prostaglandins Leukotr Essential Fatty
Acids, Vol. 48, pp 101-104, 1993.
- Horrobin DF. The role of essential fatty acids and
prostaglandins in the premenstrual syndrome. Journal
of Reproductive Medicine, Vol.28, pp 465-468,
1983.
- Korzekwa MI, Steiner M. Premenstrual Syndromes.
Clinical Obstetrics and Gynecology. Vol. 40(3):564-576,
1997.
- Kruger M.C., et al. Calcium, gamma-linolenic acid
and eicosapentaenoic acid supplementation in senile
osteoporosis. Aging Clin. Exp. Res, Vol 10,
No. 5, pp. 385-94, 1998.
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Brain
Function
- Anderson G., et al. Docosahexaenoic acid is the
preferred dietary n-3 fatty acid for the development
of the brain and retina. Pediatric Reserch.
Vol. 27 (1), pp 89-97, 1990.
- Hamazaki T et al. Administration of Docosahexaenoic
Acid Influences Behaviour of Plasma Catecholamine
Levels at Times of Psychological Stress. Lipids,
Vol. 34 (Supplement), 1999.
- Hibbein J. Fish consumption and major depression.
Lancet. Vol. 351, pp 1213, 1998.
- Kalmij S. et al. Polyunsaturated Fatty Acids, Antioxidants,
and Cognitive Function in Very Old Men. American
Journal of Epidemiology Vol. 145. No 1 pp 33-41.
- Okiniwa Y. et al. A high linolenate and high alpha-linolenate
diet induced changes in learning behaviour of rats.
Effects of shift in diets and reversal of training
stimuli. Biological and Pharmaceutical Bulletin,
Vol. 19(4), pp 536-40, 1996.
- Stoll A.L., et al. Omega 3 Fatty Acids in Bipolar
Disorder. A Preliminary Double-blind, Placebo-Controlled
Trial. Archives of General Psychiatry. Vol.
56, pp 407-412, 1999.
- Willatts P., et al. Effect of long-chain polyunsaturated
fatty acids in infant formula on problem solving at
10 months of age. Lancet. Vol. 352, pp 688-691,
1998.
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Cancer
- Begin ME et al. Differential killing of human carcinoma
cells supplemented with n-3 and n-6 polyunsaturated
fatty acids. Journal of the National Cancer Institute.
Vol 77, pp. 1053-61, 1986.
- Begin ME et al. Selective killing of human cancer
cells by polyunsaturated fatty acids. Prostaglandins.
Vol 19, pp 177-186, 1985.
- Hursting S.D., et al. Types of dietary fat and the
incidence of cancer at five sites. Prevention Medicine.
Vol. 19, pp 242 - 253, 1990.
- Kenny F., et.al. Gamma Linolenic Acid with Tamoxifen
as Primary Therapy in Breast Cancer. International
Journal of Cancer, Vol. 85, pp 643-648, 2000.
- Kokoglu et al. Profiles of the fatty acids in the
plasma membrane of human brain tumors. Cancer Biochemistry
Biophysics, Vol. 16(4), pp 301-12, 1998.
- Onogi N. et al. Suppressing effect of perilla oil
on azoxymethane-induced foci of colonic aberrant crypts
in rats. Carcinogenesis, Vol. 17(6), pp 1291-6,
1996.
- Thompson LU. Flaxseed and its lignan and oil components
reduce mammary tumor growth at late stage of carcinogenesis.
Carcinogenesis, Vol. 17(6), pp 1373-6, 1996.
- Yan L. et al. Dietary flaxseed supplementation and
experimental metastasis of melanoma cells in mice.
Cancer Letters. Vol. 124(2), pp 181-6, 1998.
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Other
conditions
- Bates D., et al. A double-blind controlled trial
of long chain n-3 polyunsaturated fatty acids in the
treatment of multiple sclerosis. Vol. 52(1), pp18-22,
1989.
- Behan P.O. et al. Effect of high doses of essential
fatty acids on the postviral fatigue syndrome. Acta
Neurologica Scandinavica, Vol. 82(3), pp 209-16,
1990.
- Belluzzi, A., et al. Effect of an enteric-coated
fish-oil preparation on relapses in Crohn's disease.
New England Journal of Medicine. Vol. 334,
pp 1557-1560, 1996.
- Bennet W.M., et al. Treatment of IgA nephropathy
with eicosapentanoic acid (EPA): a two year prospective
trial. Clinical Nephrology. Vol. 31, pp 128-131,
1989.
- Christophe A at al. Intake of Alpha Linolenic Acid,
but not of Docosahexaenoic Acid, correlates positively
with DHA content, and with parameters for fertilization
potential of human spermatozoa. Abstract for 90th
AOCS Annual Meeting, Orlando, Florida 1999.
- Clark WF, et al. Flaxseed: a Potential Treatment
for Lupus Nephritis. Kidney International.
Vol. 48(2), pp 475-80, 1995.
- Fung S.M. et al. Fish oil therapy in IgA Nephropathy.
Annals of Pharmacotherapy. Vol. 31, pp 112-115,
1997.
- Walton A.J., et al. Dietary fish oil and the severity
of symptoms in patients with systemic lupus erythematosus.
Annals of the Rheumatic Diseases. Vol. 50 (7),
pp 467-466, 1991.
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Pet Health
- Bond R, Lloyd DH. Double-blind Comparison of an
Evening Primrose Oil and Fish Oil Combination in the
Management of Canine Atophy. Veterinary Dermatology.
Vol. 4, pp 185-189,1993.
- Lloyd D, Thomsett LR. Essential Fatty Acid Supplementation
in the Treatment of Canine Atopy: A Preliminary Study.
Veterinary Dermatology. Vol. 1, pp 41-44, 1989.
- Scott DW, Miller WH, Reinhart GA, Mohammed HO, Bagladi
MS. Effect of an Omega-3/Omega-6 Fatty Acid-Containing
Commercial Lamb and Rice Diet on Pruritus in Atopic
Dogs: Results of a Single-Blinded Study. Canadian
Journal of Veterinary Research. Vol. 61, pp 145-153,
1997.
- Watson TDG. Diet and skin disease in dogs and cats.
Journal of Nutrition. Vol. 128 (Supplement),
pp 2783-2789, 1998.
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